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We have seen that Ward (1892) had no difficulty in isolating and identifying both yeasts and bacteria and establishing their role in the mixed culture' of the ginger-beer plant. The progress in understanding the role of microorganisms in the natural cycles of the elements also depended on identifying and growing the organisms concerned in these activities in pure cultures. Nitrifiers The importance of maintaining soil fertility was recognized long before the presence of microorganisms in the soil was known.
All attempts to obtain cultures of the nitrifying organisms were unsuccessful until Winogradsky thought it might be helpful to omit an organic source of nitrogen. The first isolates were made in liquid medium, but Winogradsky (1890) succeeded in obtaining colonies on a solid medium which contained "silica jelly" instead of gelatin and with ammonium chloride as the nitrogen source. This was the first isolation of Nitrosomonas and Nitrobacter. Nitrogen Fixation Pasteur's observations on the oxidative and fermentative activities of microorganisms stimulated research on soil problems.
The conclusion from this type of work was that the inheritance of acquired characters could be observed in bacteria. J ust as bacteria were the last group to be liberated from spontaneous generation, they became the last to be liberated from Lamarckian theories. Yet, within a few more years, a very different kind of acquired inheritance was identified with the discovery of the transmissible drug resistance plasmids, and, later, population studies on bacteria became very rewarding. With the development ofbacterial genetics, it became possible to examine the mechanism of enzyme induction in a new way.